::'''L70''': ''L. coryniformis''
::'''L14, L18, L22, L29, L88, L92''': ''L. Brevis''
Hop tolerance can also be encouraged with the availability of organic acids that are used by ''Lactobacillus'' for ATP (energy). [https://www.tandfonline.com/doi/full/10.1080/03610470.2020.1864710?scroll=top&needAccess=true Zheng et al. (2021)] found that many species of ''Lactobacillus'' in MRS media containing iso-alpha acids have a higher growth rate when certain organic acids are also present. In particular, pyruvate allowed for more growth, and to a lesser extent so did other organic acids such as oxalic, tartaric, and ketoglutaic acids. Citric acid, fumaric acid, and acetic acid did not have this positive effect, and acetic acid had a slightly negative effect on cell growth. Without the presence of iso-alpha acids in the MRS media, the additions of the organic acids made no difference, demonstrating that the presence of organic acids is particularly important when ''Lactobacillus'' is exposed to iso-alpha acids <ref>[https://www.tandfonline.com/doi/full/10.1080/03610470.2020.1864710?scroll=top&needAccess=true Feiyun Zheng, Tianmu Wang, Chengtuo Niu, Yun Jia, Ruilong Zheng, Chunfeng Liu, Jinjing Wang & Qi Li (2021) Proteomic Analysis of Hop Bitter Compound Iso-α-acid Tolerance in Beer Spoilage Lactobacillus casei 2-9-5, Journal of the American Society of Brewing Chemists, DOI: 10.1080/03610470.2020.1864710.]</ref>.
See also: